2010–today Director, Experimental Imaging Center, Ospedale San Raffaele, Milan
2017-todaly Full Professor of Anatomy, University Vita-Salute San Raffaele, Medical School, Italy
2000-today Full Professor of Anatomy, University of Genoa, Medical School, Italy
2002-2010 Director and co-founder, MicroSCoBiO Research Center, IFOM, University of Genoa
1992-2000 Associate Professor of Anatomy, University of Genoa, Medical School, Italy
1992-2001 Chief research unit, Department Experimental Medicine, University of Genoa
1990-1991 Marie Curie Senior Scientist, École Normale Supérieure, Paris
1985-1990 Staff Scientist, National Cancer Institute, Genoa (I)
1989 EMBO Fellow, EMBL, Heidelberg
1983-1986 Postdoc, National Cancer Institute, Bethesda, MD (USA)
1981 Training student, National Cancer Institute, Bethesda, MD (USA)
1980 Training student, Weizmann Institute of Science, Rehovot (IL)
1979-1982 Training student, Institute of General Pathology, Medical School, Federico II University, Naples (I)

1990 Commission of the European Communities Training Grant in Cancer Research Fellowship
1989 EMBO Fellowship
1984 Fogarty International Fellowship award
1983 Fogarty International Fellowship award
1982 Fogarty International Fellowship award

1987 Board in Oncology, cum laude, Federico II University, Naples, Italy
1982 Medical Degree, cum laude, Federico II University, Naples, Italy