1993 Ph.D. in Human Oncology, University of Torino, School of Medicine
1989 Medical Doctor, “Summa cum laude” and Special Award, University of Torino (Italy), School of Medicine

2007-present: Full Professor of Human Anatomy, Università Vita-Salute San Raffaele, Milano, Italy
2005-2010 Group Leader, IFOM, FIRC Institute of Molecular Oncology, Milano, Italy
2001- 2007 Associate Professor of Human Anatomy, Università Vita-Salute San Raffaele, Milano, Italy
1997-2004 Teaching activity at the PhD Program of Cell Biology – YaleUniversity, New Haven, CT, USA
1995-2005 Visiting Scientist, Department of Cell Biology and Howard Hughes Medical Institute (laboratory of Dr. Pietro De Camilli), YaleUniversity, New Haven, CT, USA
1994-2001 Assistant Professor, Department of Medical Sciences, Università del Piemonte Orientale “A. Avogadro”.
1992-1993 Maitre Assistant, Département de Pathologie, Centre Médicale Universitaire, Faculté de Médecine, Université de Genève.
1990-1991 Post-doctoral Research Fellow, Laboratory of Dr. Giulio Gabbiani, Département de Pathologie, Université de Genève, Suisse.